The submitted project represents a comprehensive palaeobotanical application of X-ray and neutron radiography. These methods, the cutting edge technologies in palaeontological research worldwide, enable the investigation of anatomical characters of fossils in a rogressive, nondestructive way. The chosen approach is particularly suitable for unique, delicate and rare specimens. Radiographic non-destructive imaging methods will provide data suitable for 3D modeling. The examination of anatomical characters of well preserved Cretaceous fossil plants will enable a better understanding of evolution of particular plant groups. The developed specific methodologies will be tested through case studies based on Cretaceous fossil plants from the pteridophyte families, conifers and angiosperms. The interpretation of newly identified evolutionary significant characters published in high impacted journals is the major results of the project.